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| 2.1 Introduction
2.2 Vegetation Sampling
2.3 Data Storage
2.4 Vegetation Data Analysis
2.5 GIS Modelling of the Vegetation Map Units
2.6 Comparisons of map units to other vegetation classifications
2. Method
2.1 Introduction
The approach involved the completion of several clearly defined
stages. These were to collate all available floristic site data;
* complete new survey work in unsampled environments;
* derive vegetation communities using statistical agglomeration
techniques;
* extrapolate the derived ecosystems across all land tenures across
the sub region;
* produce a map and report which identifies the variation and location
of vegetation communities across the sub region;
2.2 Vegetation Sampling
2.2.1 Data Audit
An audit of all systematic vegetation survey data was undertaken.
Systematic vegetation survey sites needed to conform to the following
characteristics:
* a fixed plot size within which an inventory of all vascular plant
species recorded;
* a measure of relative abundance for each species is recorded at
each site; and,
* an accurate location reference (Australian Map Grid) to within
100 metres.
As highlighted by Keith & Bedward (1999) it is possible to use
data without abundance records however this would require reducing
all other data to this level. Cover abundance scores are important
indicators when attempting to define patterns in vegetation (Keith
& Bedward, 1999). For this reason they were considered a requirement
for data used in the analysis.
A total of 780 existing systematic vegetation sites were available
for use in this project. Most sites have been extracted from systematic
surveys completed within conservation reserves and state forests
over the last 10 years. More recent regional surveys have provided
systematic data across a range of different tenures (NEFBS 1992,
CRA, 1998). The source of all site data used is shown in Table 1.1.
All site floristics and attribute data (where collected) are now
stored in the NPWS Flora Survey database.
Modifications to the existing site data were limited to a taxonomic
review and a conversion of relative abundance scores used by Thomas
(1998), Clarke and Benson (1986) and Clarke and Benson (1988) to
a six point Braun-Blanquet index. Table 2.1 provides the conversion
table for these sites.
TABLE 2.1: RELATIVE ABUNDANCE
CONVERSION SCORES
Conversion of relative abundance scores used by Thomas (1998) to the
six Point Braun-Blanquet index is as follows:
|
Code |
Relative Abundance |
Braun-Blanquet Score |
|
V: Very common |
very common, usually single spp.
dominant |
4 |
| C: Common |
common, dominance is shared by
2 to 3 other spp |
3 |
|
F: Frequent |
frequent, a spp not sharing dominance
but remains significant to the composition of the site |
1 or 3 depending on spp. |
| O: Occasional |
occasional occurrence |
1 or 2 depending on spp. |
|
R: Rare |
rare, individuals infrequently
seen, low count and crown cover |
1 or 2 depending on spp. |
|

|
Relative abundance scores used by Clarke
(1986) in the Popran NP data have been converted according to the
following table:
|
Code |
Relative Abundance |
Braun-Blanquet Score |
|
+ or no score |
For a species present but in
very low numbers |
1 or 2 depending on spp. |
| 1 |
Low numbers |
3 |
|
2 |
Moderately common |
4 |
| 3 |
Common to very common |
5 |
|
4 |
Rarely used, indicating almost
total abundance |
6 |
|
|
Clarke and BensonÕs data from Dharug
(1986) was recorded in sub quadrats nested within the 20 x 20m plots.
The presence data available from the original field sheets was converted
to a six point Braun-Blanquet index.
Table 2.2: Vegetation data
sets
|
Reference |
Location surveyed |
Data Custodian |
No. of samples in Study Area |
Species recorded |
Plot size (ha) |
Abundance measure |
|
Bell 1997 |
Tomaree National Park |
NPWS |
35 |
All vascular |
0.04 |
Braun-Blanquet |
| Bell 1997 |
Vales Point Power Station, Pacific
Power |
NPWS |
39 |
All vascular |
0.04 |
Braun-Blanquet |
|
Bell 1998 |
Glenrock SRA
Awabakal SRA
Lake Macquarie SRA
Pulbah Island NR |
NPWS |
102 |
All vascular |
0.04 |
Braun-Blanquet |
| Bell 1998 |
Popran NP |
NPWS |
20 |
All vascular |
0.04 |
Braun-Blanquet |
|
Binns 1996 |
SFNSW Morriset Management Area |
SFNSW |
116 |
All vascular |
0.1 |
Braun-Blanquet |
| Biosis 1998 |
Pinney Beach |
Lake Macquarie Council |
35 |
All vascular |
0.04 |
Braun-Blanquet |
|
Clarke & Benson 1986 |
Dharug NP |
RBG |
44 |
All vascular |
0.04 |
Relative abundance |
| Clarke 1986 |
Popran NP |
RBG |
41 |
All vascular |
0.04 |
Relative abundance |
|
Clements 1999 |
Wyong |
Wyong Council |
27 |
All vascular |
0.04 |
Braun-Blanquet |
| NPWS 1995b |
Warnervale- Site Assessment for
proposed Warnervale airport |
NPWS |
9 |
All vascular |
0.04 |
Braun-Blanquet |
|
NPWS 1994 |
NEFBS Northern Hunter Valley |
NPWS |
6 |
All vascular |
0.1 |
Braun-Blanquet |
| NPWS 1999a |
Lower North East Hunter Valley |
RACAC |
21 |
All vascular |
0.1 |
Braun-Blanquet |
|
NPWS 1999b |
Lower North East
Hunter Valley CRA |
RACAC |
127 |
All vascular |
0.04 |
Braun-Blanquet |
| NPWS1999c |
Hunter Valley- Persoonia North
Rothbury |
NPWS |
22 |
All vascular |
0.04 |
Braun-Blanquet |
|
Payne 1994 |
Munmorah SRA |
NPWS |
77 |
All vascular |
0.04 |
Braun-Blanquet |
| Payne 1998 |
Various |
Robert Payne |
27 |
All vascular |
0.04 |
Braun-Blanquet |
|
Sanders et al 1988 |
Yengo National Park |
NPWS |
30 |
All vascular |
0.04 |
Braun-Blanquet |
| Thomas 1998 |
Singleton Army Base |
Singleton Army Base |
2 |
All vascular |
|
Relative abundance |
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2.2.2 Use of External Data Sources
External data listed in Tables 2.2: Vegetation Data Sets and Table
2.6: Classifications Used for Identifying Extant Vegetation Cover
have granted permission from the relevant data custodian. This data
has kindly been provided for use in this project only.
2.2.3 Environmental Stratification for Site selection
The aim of the stratification in this process is to provide a structure
to support the project goals by ensuring the efficient allocation
of field survey effort. The focus of this study has been to gather
information on private land to establish a solid data resource to
support future data acquisition and analysis. To this end, the approach
sought to sample the widest variation in plant communities present
in the study area while having sufficient replication to explain
them. A key component was to visit plant communities that were less
likely to receive survey effort in the near future, so that a complete
picture of all communities could emerge.
A field survey program was planned in order to sample major gaps
in site coverage across the study area. Development of the stratification
and sampling regime needed to consider existing data, recent CRA
mapping, long-term data objectives and financial resources. Funds
were allocated for approximately 300 new sites, which, in combination
with available sites provided for a target of over 1100 sites in
the study area.
To efficiently sample, the location of potential variations in
vegetation communities is needed. Factors that are most likely to
affect plant distribution and abundance are moisture, nutrients
and solar energy and fire although this is not easily mapped (NPWS,
1997). Surrogates can be used to describe these factors using rainfall
distribution maps, soil maps and aspect models. Features within
these digital layers can be combined to form parcels of the landscape,
which comprise similar environmental characteristics (Neldner, 1995;
Margules & Redhead, 1995). These parcels or ÒstrataÓ are then assumed
to represent features that are likely to support similar assemblages
of plant species.
Recent CRA work completed surveys across a large area from the
Hawkesbury River to the Queensland border. Surveys focused primarily
on public land to provide information to address the forestry reform
process. Within the Hunter Sub-region (Hawkesbury to the Hunter
River) a stratification system had already been established to sample
variation in environmental characteristics across a much larger
area than that of the LHCCREMS study area. In that analysis five
data layers were used in the ARC View GIS system to highlight potential
environmental variations. These were rainfall, temperature, dominant
lithology, aspect and broad forest structure (NPWS, 1999b). As the
LHCCREMS study area represents a subset of this study area, portions
of the landscape already had some survey effort within them.
Two environmental factors were selected for stratification in this
project: soil fertility and aspect. As previous survey effort had
captured the major variations in elevation and rainfall it was considered
that soil fertility and aspect would capture the widest variation
of communities, particularly if sites were well spaced across the
region.
Soil landscapes were used as a surrogate for soil fertility. They
were chosen for three reasons. Firstly they form the most recent
and detailed mapped layer available, which describe variations in
substrate. Secondly, they inherently account for variations in topography
such as slope. Thirdly they are major recognisable land management
resource that are widely available to the community and councils.
Aspect was used to review local variations in vegetation that may
arise from the influences of shelter and exposure. Four aspect classes
were used: 0: flat, 1: 301-30¡, 2: 2-211¡, 3: 31-120¡, 4: 121 -
210¡. ARCVIEW GIS was used to combine the above aspect classes with
each soil landscape. Not all landscapes were merged with an aspect
class. Landscapes that contained slopes less than three degrees
were considered to be flat. Typically these areas contained mangrove
and wetland environments. A total of 325 strata were derived.
A proportional sampling strategy was used to identify sampling
priorities for each stratum. Sampling was initially directed toward
soil landscapes greater than 50 hectares. The proportional sampling
strategy (Sanders et al, 1988; Keith and Sanders, 1990; Keith &
Bedward, 1999) allocated sampling effort using the extant vegetation
remaining on each soil landscape in relation to the total vegetation
cover present. As an example if the Beresfield soil landscape comprised
10% of the vegetated landscape then 10% of the total number of sites
expected would be allocated to that strata.
Progress on the sampling performance was reviewed regularly. These
reviews also considered the number of sites allocated in proportion
to the uncleared area of each stratum to ensure that heavily cleared
strata received adequate sampling wherever possible.
2.2.4 Site Selection and Access
All existing survey plots were compiled and mapped across the strata
(soil landscapes and aspect) in order to identify sampling adequacy.
The position of sites was cross-referenced against soil landscape
and aspect and tabulated. Deficits in the sampling of each strata
highlighted areas requiring new survey. The majority of existing
data was obtained from public tenures or land in the reserve system.
Therefore targeting of private land was priority in this survey
to minimise sampling gaps across these areas and reduce potential
tenure bias in the classification of vegetation.
Following the identification of these areas, maps indicating survey
priorities within each local government area were produced. Each
council was visited to identify land ownership details for priority
areas. First survey priorities were those areas containing several
target strata within single owner properties. This minimised travel
time between survey sites and reduced time consuming access negotiations.
Where owners were identified, each was assigned a code. The RBCS
team arranged access to these properties. Letters were sent requesting
permission to survey. In most instances this was followed up by
a phone call. Where access was approved the area was marked for
survey. Acknowledgment goes toward all private landholders that
cooperated in the survey.
2.2.5 Field survey
The field survey collected data using standard NPWS field proformas.
Field plots were fixed to 0.04 hectares based on 20m x 20m quadrats.
Modifications were made occasionally to the plot size (usually 10
X 40 m) to match patterns in vegetation. This was applied to features
such as riparian vegetation. This quadrat size is the standard used
by NPWS and the Sydney Royal Botanic Gardens (NPWS, 1997). Over
80% of the existing plots in the region conform to this size and
it was considered both consistent and time efficient to maintain
this plot size in the region. Descriptions of field data collected
are described in more detail elsewhere (NPWS, 1995 and NPWS, 1999b).
Three field teams were established to undertake five-day field
trips to survey unsampled or insufficiently sampled strata. Each
team consisted of one experienced botanist and an assistant. New
plot data was collected between March and July 1999. Strata maps
were prepared at a scale of 1: 25 000 with the matching topographic
series. These maps highlighted the soil landscape and aspect that
required sampling within the area marked for survey. Standard NPWS
Survey techniques were applied. These involve:
- Avoiding boundaries of strata to prevent the sampling of ecotone
communities
- Avoiding local disturbances such as roads, mines quarries and
other areas of gross disturbance
- Aligning the quadrat with the contour of hillsides or elongated
vegetation communities (eg. riparian) to avoid significant local
environmental gradients.
At each site standard field survey proformas were completed (see
Appendix A). Completed survey forms provided:
- 1. lists of plant species with respective cover-abundance values.
Cover-abundance values conformed to a six-point Braun-Blanquet
scale (1-<5% and uncommon, 2- <5 and common, 3- 5-25%, 4- 25-50%,
5- 50-75%, 6- 75-100%;
- estimates of the height, cover and dominant species of each
vegetation stratum;
- measurements of slope, aspect and horizon azimuths;
- parent material; and
- qualitative notes on soil moisture, texture and depth, and disturbance
history.
2.2.6 Plant Identification Species that could not be
identified in the field were recorded to the nearest possible family
or genera. These were then collected, tagged and pressed with the
appropriate site code. All major features of the specimen were collected
where possible ie. fruit and/or flower. Subsequent to the fieldwork
the botanist identified outstanding specimens. These were noted
on the site proforma. Where positive identifications could not be
made, specimens were sent to the National Herbarium for resolution.
In the event that any rare or threatened plants were recorded in
the plot a threatened plant proforma was completed (Appendix B).
Opportunistic records were also taken where a rare or threatened
plant was located on route to the site. Interesting specimens and
range extensions were also lodged at the herbarium at the botanistsÕ
discretion.
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2.3 Data Storage
2.3.1 Site Identification
For the purpose of managing existing and new field data each survey
plot was given an 8-digit alphanumerical survey identification number,
taken from a system used by Bell et al (1993). A separate
survey identification code was given to all data to distinguish
its source. Using this system enables the reader to understand basic
geographical information about the survey site.
Example:
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The first three letters ÒGSFÓ refer to the first three consonants
in the 1:25 000 map sheet name
The fourth and fifth digits Ò07Ó refer to the site number, ie.
The seventh site on this map sheet
The sixth character ÒHÓ refers to the geology using the following
categories.
N = Narrabeen sandstone
H = Hawkesbury sandstone
W = Wianamatta shale
B = Basalt
A = Alluvium
Q = Quaternary sand
P = Permian sediments
C = Carboniferous sediments
The seventh character Ò3Ó refers to the aspect using the following
categories.
1 = 67.6 ø112.5 degrees
2 = 112.6 ø 157.5 degrees
3 = 157.6 ø 202.5 degrees
4 = 202.6 ø 247.5 degrees
5 = 247.6 ø 292.5 degrees
6 = 292.6 ø 337.5 degrees
7 = 337.6 ø 22.5 degrees
8 = 22.6 ø 67.5 degrees
The eighth character ÒCÓ refers to the morphology codes as used
in the field survey proforma (see appendix C).
2.3.2 Database Entry
Field data was entered into an ACCESS database by the field assistants
involved in the survey. Database entry windows were similar to those
used for field proformas to minimise entry errors. All species recorded
are coded using the Census of Australian Vascular Plant Species (CAPS).
New species or subspecies, as identified by the Royal Botanic Gardens,
not previously listed in the CAPS were assigned new codes to the master
CAPS database. Table 2.3 Taxanomic Revisions, provides a guide to
the genera revised in the taxanomic review.
Data from other sources was transferred electronically into the
ACCESS database.
2.3.3 Data validation
On completion of the field survey program, all data was checked
for errors. A number of steps were involved as described below.
- The database has a series of checks to prevent gross errors.
These include restrictions on the amount of numbers that can be
entered into eastern and northing fields. The database was also
designed so that vegetation structure descriptions could only
be sourced from species identified from the site species list.
- A manual check of each site held in the database was compared
to the original field sheet. Grid references were checked, as
were total number of species recorded at each site. Searches for
missing cover abundance scores were also completed.
- A number of queries using a crosscheck of field data with the
GIS system were completed. These reviewed: (1) topographic sheet
of the site code against the topographic sheet name in which the
GIS records the site (2) comparison of field elevation and the
digital elevation model in the GIS and (3) comparison of tenure
marked in the field and that recorded by the GIS.
In all cases, where errors where detected, sites were reviewed
and the correct information entered.
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2.4 Vegetation Data
Analysis
2.4.1 Taxonomic Review
Data sourced for this project has been collected over a 13-year
period by numerous observers. Over this period the taxonomy of plants
has undergone continual revision. It becomes necessary to establish
a botanical nomenclature to equate species collected from different
periods. For this project, all nomenclature was reviewed and standardised
across data sets. Synonyms for the same taxon were updated to reflect
currently accepted revisions. The treatment given in Harden (1990-93)
was used as a standard with the revisions outlined in Table 2.3.
The principle outcomes of the taxonomic review follow.
- all exotic species were identified and excluded from the analysis
dataset;
- highlighted species that were likely to have been incorrectly
identified or incorrectly entered into the database. Original
field sheets were reviewed to determine the status of these species
and where data entry errors were detected changes were made to
the database. Where data entry errors were not detected species
were reviewed against existing literature. Where this indicated
them to be outside their likely range, and no confirmation had
been made, the record was deleted;
- inconsistently collected records of species containing sub-species
or varieties. In such cases, sub-species were either lumped to
species level or were assigned to a single sub-species or variant
if only one variety is present in the study area;
- species hybrids which are not recognised formally in the literature
were assigned to one or other of the species based on the predominance
of either in proximate environments; and;
- species identified to genus level only. These were deleted from
the analysis dataset.
TABLE
2.3 TAXONOMIC REVISIONS USED SINCE HARDEN (1993)
|
Family/Genus/Species |
Authority |
|
Syncarpia |
Bean (1995a) |
| Ochrosperma oligomerum |
Bean (1995b) |
|
Babingtonia |
Bean (1997a) |
| Baeckea |
Bean (1997b) |
|
Restoniaceae |
Briggs & Johnson (1998a): Briggs
& Johnson (1998b) |
| Bursaria |
Cayzer, Crisp & Telford (1999) |
|
Prostanthera cryptandroides |
Conn (1999) |
| Boronia |
Duretto (1999) |
|
Grevillea buxifolia complex |
Hart & Henwood (1996) |
| Corymbia |
Hill & Johnson (1995) |
|
Angophora inopina |
Hill (1997) |
| Phyllanthus gastroemii |
Hunter & Bruhl (1997) |
|
Brachyloma daphnoides ssp.
daphnoides |
Hunter & Williams (1994) |
| Stipa & Austrostipa |
Jacobs & Everett (1996) |
|
Danthonia |
Linder (1997) |
| Notodanthonia |
Linder & Verboom (1996) |
|
Acacia linearifolia |
Maslin (1994) |
| Grevillea obtusiflora |
Olde & Marriott (1994) |
|
Lissanthe strigosa |
Powell & Wiecek (1994) |
| Goodia lotifolia |
Ross (1997) |
|
Pomaderris |
Walsh & Coates (1997) |
| Persoonia pauciflora |
Weston (1999) |
|
Eriostemon |
Wilson (1998a) |
| Macrozamia flexuosa |
Hill (1998) |
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2.4.2 Classification Analyses
The use of semi quantitative field survey methods allows the application
of numerical techniques to assist with the identification of vegetation
communities. Numerical classification techniques are not an end
themselves but a means to review and compare surveyor field experiences
with raw data. In this way, biases and patterns easily overlooked
in the field are avoided. Such techniques are aimed at present hypotheses
to help describe patterns in the field. Whether these are accepted
rejected or modified remains a subjective element of the chosen
classification system. However, the benefits of such an approach
are that an explicit and repeatable method is employed.
The PATN analysis package (Belbin 1994) was used to group sites
based on the similarities of species occurrence and abundance between
sites. PATN provides a range of modules and algorithms from which
to undertake analyses of ecological data. ASO, FUSE, GDEF and DEND
were used to establish the association matrix for hierarchical classification,
clustering and the production of a dendrogram.
PATN analyses were performed on 2360 sites drawn from the wider
Hunter Region to include the Upper Hunter region, Wollemi and Yengo
National Parks. Of these, 1117 sites fell within the LHCCREMS study
area. The pooling of all available site data in the region enabled
site data occuring on the edge of the study area to be placed in
a broader regional context.
Several different analyses were performed to identify sensitivities
in the data. Bray-Curtis and Kulzcynski measures of association
were both tested on data using raw cover abundance codes for each
species and a simpler presence-absence data set to derive dissimilarity
scores between sites. These scores are calculated by comparing all
possible pairs of sites in the data set (eg. Site1 and Site 2, Site
1 and 3 and so on.). A large (association) matrix describing the
dissimilarity of the pairs is produced. An unweighted pair-group
arithmetic averaging (UPGMA) clustering strategy was applied to
the matrix to derive a hierarchical classification. The default
beta value of ø0.1 was used for all analyses.
Dendrograms were derived to show the degree of dissimilarity between
individual sites and groups of sites. The review of groups commenced
at the highest levels of the dissimilarity scale. The dendrograms
provided a coarse overview of the way in which the sites were positioned
relative to one another.
To commence the fine scale analyses of data, homogeneity analyses
(Bedward et al., 1992) was applied to the association matrix.
This technique provides a useful guide to the benefits of improvements
in species homogeneity for increasing numbers of groups or clusters
made from the data set. If every single site in the dataset were
made a group or Community, its characteristics would be perfectly
homogenous. This would mean 2360 groups. At the other end of the
scale, using only two groups would result in large within group
variation so homogeneity would be low. The aim is to select the
number of groups appropriate to the dataset to maximise returns
to group homogeneity while minimising the number of groups used
to describe the patterns in the data
Seventy groups were initially used as a means of interrogating
the data. Group definition files were extracted, and species characteristics
of each group were produced. A check for misclassified sites within
the group was undertaken using 'GDFChk'. This reviews the
grouping and highlights misclassified sites by examining the characteristics
of the neighbouring sites in the group. Where identified, sites
were reviewed and either withheld from further analyses, reallocated
to a new group or kept with the existing group.
Groups were successfully split, where such splits reinforced sub
groups based on physical characteristics of sites and floristic
composition. Most often splits were made where distinct changes
in substrate characteristics were apparent. Further, groups were
refined at lower levels in the dissimilarity index where such splits
improved the definition and recognition of floristic composition.
No further splits were made when patterns presented did not accord
with field experience or where features were unlikely to be mappable.
A number of core groups were readily apparent using these methods
and required no further analyses. Other groupings required individual
review of sites to determine the reasons behind clustering.
2.4.3 Vegetation Map Unit Descriptions
Each Map Unit has been described using data from 20 X 20 metre windows
into the vegetation characteristics of the study area. These windows
or plots can be used to characterise the species composition of the
Map Unit. Clustering of plots in the analyses can indicate that combinations
of species are repeating themselves in the landscape. In the study
area importance of a species or groups of species to defining a Map
Unit not only hinges on its abundance within the sites used to make
up the group, but amongst sites used to describe all other Map Units
defined from the data set.
A software program known as FIDEL was developed for use in the
vegetation mapping of the South East Forests (Keith and Bedward;
1999). This program allows relevant data to be extracted from the
sites that make up the defined Map Units to indicate which species
are driving the distinction between one Map Unit and another. Keith
and Bedward (1999) termed these diagnostic species.
In some Map Units a species is dominant both in terms of its frequency
of detection at sites and its abundance while in other Map Units
it occurs patchily, at low abundances or not at all. This can be
quantified by setting user defined rules to determine diagnostic
species.
A table mirroring that used by Keith and Bedward (1999) is presented
here. It should be noted however that the criteria for defining
positive species has been amended to allow for greater number of
locally abundant species to be included as positive diagnostics.
TABLE
2.4: Definitions of diagnostic species
| Occurrence of
Species within Target Map Unit |
ÊÊ |
Occurrence of Species in Residual Map Units |
|
Frequency >50% AND C/A >2 |
Frequency <50% OR C/A <2 |
Frequency =0 |
|
Frequency <35% AND C/A >2 |
Uninformative |
Positive diagnostic |
Positive diagnostic |
|
Frequency <35% OR C/A <2 |
Uninformative |
Uninformative |
Positive diagnostic |
|
Frequency =0 |
Negative diagnostic |
Uninformative |
- |
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* C/A = Cover abundance
Based on this table two categories have been defined. Positive
diagnostic species are those that have a higher frequency recorded
within the group and / or have been recorded at a higher median
cover abundance than in all other Map Units. Also species that are
only recorded within the target Map Unit and in no other are considered
positive. Uninformative species are those species that do not display
unique characteristics to the target Map Unit. They are likely to
be species common to many communities, or are those recorded at
low frequency and / or very low abundance.
Map Unit profiles contain all species defined as positive diagnostics.
A list of uninformative species is given for all canopy species
and species considered by the authors to assist with the presentation
of a visual picture of the Map Unit.
Summary structural data is provided for each Map Unit. The mean
upper height, the range in field heights, the mean cover abundance,
the standard deviation of the cover abundance estimates and the
number of sites used to describe each structural layer is given.
A summary species richness figure and the associated standard deviation
amongst sites in the Map Units are also provided.
A canopy label is also provided to assist those familiar with tree
species only to impart a rough guide to the type of environment
expected.
It needs to be reinforced that the presence or absence of some of
these species does not indicate that the mapping is incorrect, rather
that other non canopy species are likely to be influencing the classification.
Consultation with the full floristic profiles and on site presence
is required.
2.5. GIS Modelling of the Vegetation Map Units
2.5.1 Development of Spatial Data Layers
Spatial data layers describing the abiotic characteristics of the
study area were compiled at 25 metre square grid cell resolution
for use in vegetation modelling. These data layers were derived
by NPWS in the following way:
- Terrain variables including topographic position, roughness
index, wetness index, stream order, slope, aspect, solar radiation
indices were derived from a 25m grid digital elevation model supplied
by the NSW Land Information Centre.
- Climatic surfaces (Table 2.5) were derived using ESOCLIM (Hutchinson
1989). This provided layers describing mean annual figures as
well as coldest and hottest months and driest and wettest months.
The Bureau of Meteorology provided rainfall and temperature data.
- Dominant lithological features were grouped from soil landscape
mapping within the region (DLWC, 1992-7). Three different layers
representing soil parent material were derived. The first represented
a 14 scale class which amalgamated mapped landscapes according
to their dominant lithology (tertiary alluvium, quartz sandstone,
quaternary sand, quaternary sediments, quaternary alluvium, acid
volcanics, basic igneous, granitic rocks, sedimentary (coarse
grained) sedimentary (fine grained). Secondly, the 14 class lithology
was grouped into five classes to provide a relative index of soil
fertility. This ranged from basic igneous as the highest fertility
through to quartz sandstones at the lowest. The unique soil landscape
code provided the third substrate layer.
- Other variables were generated directly in the Arcview GIS
system to provide a grid for easting and northing values and to
generate layers indicating distance from features including streams,
geological features and the coastline.
-
TABLE 2.5:
SPATIAL DATA LAYERS USED IN MODELLING
-
| GIS COVERAGE |
DESCRIPTION |
| Altitude |
Elevation above sea level
(metres) |
| Slope |
Inclination from horizontal
(degrees) |
| Aspect |
Deviation from grid north
perpendicular to slope (degrees) |
| Aspect Index |
Categorical index of aspect
(0: flat, 1: 301-30¡, 2: 211-300¡, 3: 31-120¡, 4: 121-210¡) |
| Solar Radiation Index ø January
(Summer) |
Index representing topographic
exposure to solar radiation calculated from slope, aspect,
horizon azimuth and latitude. Varies below 100 for sheltered
sites and above 100 for exposed sites |
| Solar Radiation Index ø July
(Winter) |
Index representing topographic
exposure to solar radiation calculated from slope, aspect,
horizon azimuth and latitude. Varies below 100 for sheltered
sites and above 100 for exposed sites |
| Solar Radiation Index |
Continuous index representing
topographic exposure to solar radiation calculated from
slope, aspect, horizon azimuth and latitude. Varies below
100 for sheltered sites and above 100 for exposed sites |
| Wetness Index |
Continuous index representing
the volume of water draining to a given point in the landscape |
| Local Topographic Position
(S) |
Continuous index (0-100)
representing proportional distance between local ridge
(100) and local gully (0) |
| Neighbourhood Topographic
Position (100) |
Difference between altitude
of a central cell and mean altitude of cells within a
1 x 1 neighbourhood |
| Neighbourhood Topographic
Position (300) |
Difference between altitude
of a central cell and mean altitude of cells within a
3 x 3 neighbourhood |
| Neighbourhood Topographic
Position (500) |
Difference between the altitude
of a central cell and mean altitude of cells within a
5 x 5 neighbourhood |
| Neighbourhood Topographic
Position (700) |
Difference between the altitude
of a central cell and mean altitude of cells within a
7 x 7 neighbourhood |
| Neighbourhood Topographic
Position (900) |
Difference between the altitude
of a central cell and mean altitude of cells within a
9 x 9 neighbourhood |
| Neighbourhood Topographic
Roughness (100) |
Standard deviation of altitude
within a neighbourhood
of 1 x 1 cells |
| Neighbourhood Topographic
Roughness (300) |
Standard deviation of altitude
within a neighbourhood
of 3 x 3 cells |
| Neighbourhood Topographic
Roughness (500) |
Standard deviation of altitude
within a neighbourhood
of 5 x 5 cells |
| Neighbourhood Topographic
Roughness (700) |
Standard deviation of altitude
within a neighbourhood
of 7 x 7 cells |
| Neighbourhood Topographic
Roughness (900) |
Standard deviation of altitude
within a neighbourhood
of 9 x 9 cells |
| Annual Rainfall |
Mean total yearly rainfall
(mm) |
| Rainfall of Wettest Month |
Maximum mean monthly rainfall
(mm) |
| Rainfall of Driest Month |
Minimum mean monthly rainfall
(mm) |
| Minimum Temperature of Coldest
Month |
Mean minimum monthly temperature
(¡C) |
| Maximum Temperature of Hottest
Month |
Mean maximum monthly temperature
(¡C) |
| 5-class Parent Material |
Major Fertility Classes based
on amalgamation of geologies (Low Fertility- Hawkesbury
Sandstone, Quartz sands to High Fertility -Basic Igneous) |
| 9-class Parent Material |
Dominant lithologyÕs |
| 393-class Parent Material |
Raw Soil Landscape Codes |
| Vegetation Structure |
Vegetation Structure was
derived from all available digital mapping in the study
area. Classifications were converted to standardise with
the 17-class structure used by DLWC. |
| Forest Types |
Types and mosaics interpreted
from aerial photos according to Baur (1989). All vegetation
mapping using canopy species descriptors was transcribed
to this classification system. |
| Distance from Coast |
Shortest distance from coast
(metres) |
| Easting |
Australian map grid |
| Northing |
Australian map grid |
| Distance from Basalts |
Shortest distance from Basalt
soils (metres) |
| Distance from Fine Sediments |
Shortest distance from Fine
sediment soils (metres) |
| Distance from Narrabeen |
Shortest distance from Narrabeen
sandstone soils (metres) |
| Distance from stream order
ø stream order 1 |
Shortest distance from stream
order 1 |
| Distance from stream order
ø stream order 1-2 |
Shortest distance from stream
orders 1 and 2 |
| Distance from stream order
ø stream order 1-3 |
Shortest distance from stream
orders 1 to 3 |
| Distance from stream order
ø stream order 2-4 |
Shortest distance from stream
orders 2 to 4 |
| Distance from stream order
ø stream order 4-9 |
Shortest distance from stream
orders 4 to 9 |
| Distance from stream order
ø all orders |
Shortest distance from all
stream orders |
|
|
2.5.2 Deriving the Extant Vegetation Layer
Mapping of vegetation cover was not available as a consistent data
layer across the study area. Sixteen data layers described in Table
2.6 were merged together in ArcView to provide a single coverage
to a scale of at least 1:25 000. Some mapping has been captured
from larger scale aerial photography and these have been noted in
the table where known. Considerable variation is apparent between
each of the coverages in terms of geo-referencing accuracy and attributes
collected. The most notable discrepancies will emerge between criteria
used to map remnant vegetation. Some mapping may delineate remnants
less than five hectares others will not map under this size and
hence small patches of vegetation may be excluded.
Mapping completed by DLWC (1999) covers a wide range of structural
types, native and exotic vegetation by using canopy cover estimates
and understorey features. Extant vegetation was delineated by any
forest or woodland structure that supported a canopy cover greater
than 10%. All mapped areas of Rainforest, Littoral rainforest, Mangrove,
Heath and Swamp were included. Areas described as No Mature Trees,
Scattered Trees or Trees in Clumps were only included where canopy
cover was greater than 10% and contained a visible understorey.
Scale and accuracy of attributes in each of the coverages was unknown.
Attributes relating to either vegetation classification or vegetation
structure were retained for use in deriving the map of vegetation
Map Units (see 2.5.3).
No aerial photo interpretation is available over large areas of
Yengo National Park west of the Wollombi Valley. Extant vegetation
has been derived for this area using map interpretation of Landsat
images taken in 1989 (NPWS, 1990).
No information on the condition of vegetation has been consistently
mapped or described in the study area.
TABLE 2.6:
DATA LAYERS USED FOR DERIVING EXTANT VEGETATION COVER
|
DATA LAYER |
MAPPER |
CUSTODIAN |
YEAR AND SCALE OF PHOTO |
CRITERIA |
COVERGAE EXTENT |
|
Structural Vegetation Mapping |
unknown, various |
DLWC |
1:25000, various |
Remnants <5 ha |
LGA' s of Newcastle, Cessnock,
Maitland and Gosford |
| Vegetation Communities of Wyong
LGA |
R. Payne |
Wyong Council |
1994, 1:25000 |
Unknown |
Wyong LGA, excluding land managed
by SFNSW |
|
Vegetation Communities of Port
Stephens LGA |
R. Payne |
Australian Koala Foundation |
1994, 1:25000 |
Unknown |
Port Stephens LGA |
| Vegetation Communities of Lake
Macquarie LGA |
Biosis |
Lake Macquarie Council |
1994, 1:4000 |
Forest Communities excluding
wetlands |
Lake Macquarie LGA |
|
Wetland Communities of Lake Macquarie |
Shortland Wetland Centre |
Lake Macquarie Council |
1998, 1:4000 |
Wetlands of Lake Macquarie -
All wetlands protected under SEPP 14, all identified by previous
mapping and others greater than 1-2 Ha. |
Lake Macquarie LGA |
| Forest Types of State Forests,
Morisset Management Area |
1970 - 1995, various |
SFNSW- Morisett Forestry District |
1:16000, 1:25000 various |
Polygons > 2ha |
State Forests of Morisset and
Buladelah Management Areas |
|
Vegetation Communities of Brisbane
Waters |
Benson & Fallding 1981 |
NPWS |
1976, 1:25000 & reduced to 1;50000
at publishing |
Unknown |
Brisbane Waters National Park |
| Vegetation Communities of Bouddi
NP |
Mc Rae 1990 |
NPWS |
1;25000, 1980 |
Unknown |
Bouddi National Park |
|
Vegetation Communities of Dharug
NP |
Clarke & Benson 1986 |
RBG |
1979, 1:16000 |
Unknown |
Dharug National Park |
| Vegetation Communities of Glenrock
& Awabakal State Recreation Areas |
Bell 1998 |
NPWS |
1:25000 Newcastle 1993, Lake
Macquarie 1994 |
Unknown |
Glenrock and Awabakal State Recreation
Areas |
|
Vegetation Communities of Tomaree
NP |
Bell 1997 |
NPWS |
1993, 1:25000 & 1992 at 1:16000 |
Unknown |
Tomaree National Park |
| Sepp 14 Wetlands |
unknown |
DUAP |
1981-1982, 1:25000 |
Wetlands delineated from 1:25000
aerial photos |
Coastal Wetlands of NSW |
|
Koorangang NR and Hexham Swamp
NR Vegetation Mapping |
Winning (1996) |
NPWS |
1992 1:25 000 |
Wetland vegetation map units |
Wetlands of the Lower Hunter
Estuary incl. Koorangang and Hexham NR. |
| Vegetation Communities of Popran
NP |
Clarke & Benson 1987 |
NPWS |
unknown |
Vegetation Communities |
Popran NP |
|
Vegetation Communities of Lake Macquarie SRA, Pulbah Island
NR |
Bell, 1998 |
NPWS |
1996, 1;16000 and 1:4000 |
Vegetation Communities |
Lake Macquarie SRA, Pulbah Island
NR |
| Broad Old Growth Mapping of State
Forest Tenure |
NPWS, 1996 |
NPWS |
1994, 1:25000 |
Rainforest delineated by areas
greater than 2 hectares with less than 30% pyrophytic vegetation
in the canopy and visible rainforest understorey |
All State Forests in Morisset
Forestry District |
|
|
2.5.3 Deriving the Vegetation Map Unit Modelling Rules
The technique used to describe the distribution of each ecosystem
in the region draws on that used by Keith and Bedward (1999) for
the Eden CRA. This technique seeks to derive statistical relationships
between sites used to define a map unit and their environmental
characteristics. In essence, a series of equations is developed
that combines selected values of the environmental data layers to
describe the distribution of floristic assemblages. However, in
most instances multiple rather then single equations or Ôdecision
rulesÕ as it is known, are used to model a Map Unit using GIS.
A new piece of interactive modelling software (ALBERO) developed
by (Keith and Bedward, 1998) was used to generate decision rules
by statistical induction and expert input. The software is a tool
to structure arguments that may describe relationships between the
location of ecosystems and spatial variables. These potential relationships,
supported by a pre-defined level of statistical significance, can
be accepted or rejected by the user. ALBERO builds a decision tree
or arms of equations to describe the distribution of all communities
in the study area. Building a decision tree is a manual one with
ALBERO presenting relationships between Map Units and data layers,
and suggesting critical thresholds that continually discriminate
one Map Unit from another by using its position in the landscape.
ALBERO relies heavily on the user knowledge of the study area.
To use ALBERO a data matrix is needed to provide a raw value for
each variable at each site. This is generated using a sites file
and an output file known as ÔEnvarsÕ. Final vegetation Map Unit
numbers are allocated to each site. At the commencement of decision
rule construction ALBERO contains a list of each Map Unit and the
number of sites used to describe them. ALBERO then searches for
significant splits using the chi square statistic to discriminate
between Map Units based on the data matrix values. Each split provides
two new branches requiring resolution. At the top of the tree all
sites appear. The aim is to reach an end point in the tree to account
for a unique environmental niche where one or more sites describing
the Map Unit fall. At the start of the tree spatial variables which
depict regional variation are used most frequently eg rainfall and
broad geology. To resolve variation between Map Units at the end
of the tree, finer local scale variables are used such as shelter
indices.
There are many combinations of rules that could be used to describe
the distribution of Map Units. The process is one of iteration,
where review and further review of derived maps is carried out.
Fine scale adjustment of rule sets can be made at any time.
2.5.4 Mapping Pre-1750 Distribution of Map Units
Where all sites and Map Units are resolved, the derived rules are
then exported and run in the GIS system to derive a map of the Map
Units. Each 25 metre grid cell is allocated to a Map Unit using
all spatial variables used in the decision rules. The derived map
then represents the pre-European distribution of all vegetation
assemblages. The map is then cleaned using a filter of communities
less than a four by four neighbourhood. That is, the data is smoothed
in outlying grids by evaluating the values in the 4 surrounding
grid cells and is amalgamated with the dominant neighbouring Map
Unit. This removes small slithers and artefacts of the intersects
between the GIS layers which are less than 0.25 hectares.
2.5.5 Mapping Extant Distribution of Map Units
Extant distribution was mapped using the results of the modelled distribution
in combination with finer scale information from existing vegetation
mapping where it was available. The resolution of the derived model
is based on soil landscape layers produced at 1:100 000 and in the
case of the Cessnock area at a scale of 1:250 000. Fine scale mapping
of vegetation communities and structures listed in Table 2.6 were
used to refine modelled outputs where necessary. A suite of rules
were used for this purpose:
- Where mapped polygons of existing vegetation were enclosed within
mapped units of the vegetation model these were directly attributed
the model value unless overstorey species contradicted the diagnostic
species of the modelled map unit. In such instances, polygons
were reviewed and ascribed an analogous map unit based on proximate
sites, field knowledge and community description. Most examples
dealt with fine scale swamp communities that were not delineated
on soil landscapes given their size, but conformed to characteristics
of neighbouring larger mapped swamp patches.
- Polygons delineating rainforest-dominated canopy were attached
to the model based on overlap with modelled rainforest øwet sclerophyll
Map Units and sites.
- Polygons defining mangroves and saltmarshes were directly assigned
to the analogous Map Unit.
- Where API delineated variations in structure within a modelled
map unit this was ascribed a structural sub unit. This was prevalent
on sandstone and quaternary sand heath-woodland complexes. This
recognises that local structural variations are present although
floristic composition is relatively constant.
- Polygons that were well sampled in existing mapping (Tomaree,
Glenrock, Awabakal and Lake Macquarie NPWS reserves) were directly
ascribed the regional map unit to which the sites conformed.
- Polygons which mapped vegetation extant only (Biosis, 1999)
or were ascribed a single structural value (DLWC, 1999) were cut
directly to the modelled map units. The modelled Map Unit of similar
structure, which comprised the greatest proportion of each polygon,
was ascribed.
- Where the modelled map unit contained a vegetation structure
similar to the mapped vegetation community but different character
species the polygon was reviewed using field knowledge and proximate
sites. Where information supported the former, the polygon was
ascribed to the model. Where the later, the polygon was given
the analogous regional map unit. Where it remained uncertain the
model map unit was accepted and attributed to the polygon.
All polygons were systematically reviewed in this way. As would
be expected some joins between each of the different extant vegetation
mapping sources are not smooth. These were resolved where ever possible
using the model. Where resolution was not possible unmatched polygons
were not resolved.
Slithers and artefacts of the GIS layers were filtered in ARC-INFO.
All polygons less than 600 m2 were merged to the largest neighbouring
polygon. To assist in removing long slithers larger than this size
an index was generated to indicate an area to perimeter ratio (area
(m2)/perimeter (m)) for each polygon. Polygons that contained index
values less than 10 were merged to the largest adjoining polygon.
2.6 Comparisons of map units to other vegetation
classifications
2.6.1 Within Rems Study Area
Comparisons were drawn between some existing classifications and
assemblages defined in LHCC REMS as an example of how communities
may relate. This was achieved by assessing floristic composition,
structural form, geological substrate and distribution. No comprehensive
review was made of all communities previously described in the study
area.
2.6.2 External to Rems Study Area
A comparison was drawn between Ecosystems derived from Forest Ecosystem
Classificationand Mapping for the Upper and Lower North East (NPWS,
1999a). This involved comparing floristic composition, frequency and
distribution of each Map Unit to Forest Ecosystems derived for the
north coast. The statistical approach used in the Upper and Lower
North East was based on the analysis of full floristic data within
previously defined groups of canopy species (Broad Canopy Classification
ø Forest Types). The analyses completed for this project did not determine
apriori what the canopy groups should be.
A match was only drawn where the majority of dominant species were
shared in the canopy, mid and lower stratum. This comparison is
limited to Eucalypt dominated forests. Some further suggestions
have been made where possible between sand and swamp communities
based on floristic data presented by Myserscough and Carolin (1986)
for adjacent the Myall Lakes area.
Profiles for each Map Unit provide an indication of the extent
of similarity with assemblages outside the region
2.6.3 Map Accuracy
FEWG (1987) recommended that all vegetation maps should be evaluated
with an explicit validation process. Keith and Bedward (1999) argue
that it is important where large scale conservation planning exercises
such the CRA use such information as a key information source. Mapping
can never perfectly represent all features in the landscape and
to this end all maps will be incorrect somewhere. The general test
is whether a sample taken describes a Community that matches that
identified on the map. Most mapping exercises do not explicitly
test for accuracy. It is generally assumed that in the case of qualitatively
derived maps the level of reliability will be proportional to the
amount of field time available and the skill of the mapper.
Two approaches were used to refine and test map accuracy during
this project. Firstly, several iterations of mapping rules were
constructed in concert with all available digital vegetation mapping
in the study area. Secondly, 54 sites used to describe the vegetation
communities were extracted and set aside during the modelling exercise.
These sites were selected from those communities that were likely
to be most extensive and therefore most difficult to map. These
sites were then placed back on the map at the completion of the
modelling process. The accuracy of the Community described by the
site to that presented on the map was tested using windows of a
100 metre and 250 metre radii.
An interim review period was implemented for the maps to be reviewed
by the seven councils. Errors in identifying extant vegetation and
model inaccuracies were highlighted. These areas were targeted in
a two-week field validation period and the model was subsequently
reviewed where problems were identified.
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|

Regional Biodiversity Conservation
Strategy
- User Guide
- Local Government
area Plant Species Lists
- Module 1 Fauna Surveys
- Comprehensive
technical report
*
Acknowledgements
*
Method
*
Results
*
Discussion
*
References
*
Appendix A
~ Survey Form
*
Appendix B
*
Appendix C
*
Appendix D
~ Map Unit Profiles
*
Appendix E
Copyright 2003, Hunter Councils Inc
as legal agent for the
Lower Hunter & Central Coast Regional Environmental Management
Strategy |
|
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